Highlights d Cities possess a consistent ''core'' set of non-human microbes d Urban microbiomes echo important features of cities and city-life d Antimicrobial resistance genes are widespread in cities d Cities contain many novel bacterial and viral species

The ancient African crop, finger millet, has broad resistance to pathogens including the 19 toxigenic fungus Fusarium graminearum. Here we report the discovery of a novel plant 20 defence mechanism, resulting from an unusual symbiosis between finger millet and a root-

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