Animal-sediment relationships of two benthic communities (Crustaceans and Polychaetes) were studied around Mandvi coast in the Gulf of Kachchh, Western India. This coast consists of many micro-geomorphic landforms in which benthic communities are inhabited and select their niches and produce endemic biogenic structures. Five intertidal subfacies have been described and four types of grounds are identified, based on substrate consistency. 18 species of crustaceans, 15 species of polychaetes and unsegmented worm nemertea have been identified. Crustacean behavioural activities were observed in dunes, beaches and ridge-runnel in the form of burrowing, pellet making, feeding and crawling traces. Pelleted wall lining burrows of the suspension feeder stomatopodean species of Oratosquilla striata are also abundant in runnels. Motile, deposit feeder polychaetes are abundant on the ridges and are occasionally found on the lower reaches of the beaches, while suspension and filter feeders are found in the runnels. Lagoons consist of mainly grouped funnel branched burrows of Oniphus eremita which is identical to ichnogenus Balanoglossites. Nemertea, which are opportunistic algal grazers, have exploited restricted niches for dwelling-feeding purposes and constructed vertical burrow with pentamerous conical mound. The shore platform consists of cemented, calcareous tubes of filter feeder Serpula along with symbiotic encrusters like Ostrea and barnacles. Ichnocoenoses are discussed and three-dimensional ichno-sedimentologic models are reconstructed for Beach, Ridge, Runnel and Lagoon of the Mandvi intertidal zone.
Infaunalization has been regarded as representing a response to increased predation pressures and is therefore central to the Mesozoic marine revolution, which gives pre-eminence to the role that enemy-directed evolution has played as a driving force of biotic change. Our ichnologic compilation from 39 Middle Triassic to Late Jurassic shallow-marine siliciclastic units allows us to evaluate the vertical partitioning of the infaunal ecospace through the application of the ichnoguild concept. This study shows that infaunal communities experienced a marked increase in ecospace utilization during the Early Jurassic, reaching a plateau during the rest of the Jurassic. This trend is expressed by an increase in the maximum number of ichnoguilds per community and per tier and in the number of ichnotaxa per ichnoguild. This pattern shows important partitioning of the infaunal ecospace into a series of tiers and that several organisms were able to exploit the same resources available at discrete sediment zones below the sea bottom. The increase in the maximum number of ichnoguilds per community and per tier suggests that niche partitioning was a key factor in a more efficient use of the infaunal ecospace and in driving alpha diversity. However, the increase in the number of ichnotaxa per ichnoguild indicates that ichnoguilds were packed with organisms exploiting similar resources, arguing against the role of competitive exclusion in structuring communities. Because several episodes of predation increase took place during the Mesozoic, an unequivocal link between predation pressures and infaunalization cannot be demonstrated empirically.
Dhosa Oolite Member of the Jumara Formation comprises alternating bands of oolitic limestones and shales, exposed in Jhura dome of Mainland Kachchh, Western India. This sequence is highly bioturbated and exhibits a moderate diversity and behaviourally complex assemblage of ichnospecies. The rhythmically bedded sequence shows three different levels of preservation of traces. Epichnial tiering consists of moderately bioturbated oolitic limestone exhibiting horizontal or low-angle protrusive/retrusive biogenic laminae, commonly dominated by feeding structures like Rhizocorallium jenense, R. irregulare, Zoophycos brianteus and Zoophycos isp. The endichnial structures within the oolitic limestone can be separated into two different preservational trace fossil suites. The endichnial shallow suites consist chiefly of deposit feeders like Chondrites intricatus, C. targionii, Planolites beverleyensis, Taenidium cameronensis, Thalassinoides isp., Z. brianteus, Z. cf circinnatus and Zoophycos isp. and few suspension feeder forms like Palaeophycus tubularis; while endichnial deep suites consist of Chondrites intricatus, Skolithos linearis and Zoophycos isp. Hypichnial structures consists abundant, cylindrical, branched, horizontal, large-sized three dimensional feeding burrows of Thalassinoides isp. and somewhat irregular, obtuse angle ramification burrows of Phycodes isp., which are attached to the lower surface of the casting medium. The trace fossil association indicates Cruziana ichnofacies and abundance of Zoophycus species below the fair weather wave base level is largely a preservational artifact. The preservational processes of the trace fossils indicate soft substrate and diversity and their abundance reflects the other palaeoecological parameters of the open shallow marine environments.
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