The St. Cassian Formation, Italy, has yielded the most diverse marine invertebrate fauna known from the Triassic. A quarter of all described Triassic gastropod species has been reported from this formation. Most of the gastropod species from the St. Cassian Formation were erected in the 19th century and many of them are known only from their original figures and descriptions. The failure to study type specimens resulted in many erroneous identifications by subsequent authors. Here, we revise the slit band gastropods (Pleurotomariida) from the St. Cassian Formation—one of the major groups present in this formation. A total of 77 nominate Pleurotomariida species belonging to 29 genera and 11 families are present in the St. Cassian Formation which comprises approximately 14 % of the total nominate gastropod species of that formation. In addition, we revise several taxa that had been wrongly assigned to Pleurotomariida. As other gastropod clades, Pleurotomariida experienced a major extinction at the end-Permian mass extinction event. As in the Late Palaeozoic, their relative abundance in gastropod faunas continued to be 30 % in some Anisian faunas but decreased to 5–10 % afterwards. Their diversification at generic level became interrupted by an extinction event within the Carnian, probably by the Carnian Pluvial Event. As a result of their sluggish recovery compared to the other gastropod groups, their species diversity decreased from 26 % during the Permian to 18 % during the Triassic. Type specimens of the following genera are studied: Proteomphalus, Rhaphistomella, Temnotropis, Kittlidiscus, Stuorella, Schizogonium, Wortheniella, Bandelium, Lancedellia, Rinaldoella, Pseudowortheniella, Paleunema, Ampezzalina, Bandelastraea, Cheilotomona, Pseudoscalites, Delphinulopsis, and Cochlearia. Nine new pleurotomariidan genera are erected: Amplitomaria, Pseudoananias, Lineacingulum, Pressulasphaera, Cancellotomaria, Acutitomaria, Lineaetomaria, Nodocingulum, and Striacingulum. Eight new species are described: Schizogonium undae, Acutitomaria kustatscherae, Wortheniella klipsteini, Wortheniella paolofedelei, Rinaldoella tornata, Nodocingulum ernstkittli, Nodocingulum? turris, and Laubella subsulcata. Eoworthenia frydai is a new replacement name for Worthenia rarissima Barrande.
Drill holes provide valuable information about palaeoecological interactions in fossil ecosystems, but the Jurassic drill‐hole record is scarce. We report circular drill holes in the infaunal bivalve Eothyasira antiqua(Münster in Goldfuss) from the Pliensbachian grey shales of the Amaltheenton Formation of Franconia, southern Germany. The outcrop of the Amaltheenton Formation at Buttenheim yields a rich bivalve fauna (57 species), which represents a typical soft‐bottom community. Eothyasira antiqua, the yet oldest known member of the family Thyasiridae, represents only 1% of the total bivalve specimens from Buttenheim, but shows an exceptionally high frequency of drill holes. With a drilling rate of 81% of the well‐preserved articulated specimens, this occurence is unique compared with other drilled bivalve records from the Mesozoic and even from Recent assemblages. The drill holes resemble those of modern naticids with a larger outer opening and are assigned to the ichnospecies Oichnus paraboloides Bromley. Statistical methods show a non‐random distribution of the drill holes over the shell surface with a concentration on the central flank. The drill‐hole producer was highly efficient in drilling and exhibited a remarkable prey and site‐selective behaviour. The drill holes on the moderately deep infaunal Eothyasira suggest that the predator was also infaunal for at least parts of its life and could cope with poorly oxygenated substrate conditions occurring temporarily in the Amaltheenton environment. Drill‐hole characteristics point to a gastropod as producer. None of the about 50 gastropod taxa of the Amaltheenton Formation can be associated with the drill holes with certainty, but Hayamia reticulata (Münster) seems to be the most likely candidate. The results of this study support the opinion that drill holes are generally a scarce phenomenon in the Early Jurassic generated by rare, probably highly specialised predators. Preservational biases are considered to have been less important.
Pleurotomariida have the longest fossil record among living gastropods and are diverse and abundant in the middle and upper Palaeozoic. Its traditional classification is based on adult shell characters. The early shell morphology has been largely unknown. We describe exceptionally well‐preserved Pleurotomariida from the Pennsylvanian marine shales of Texas, Oklahoma, Kansas and Ohio. In total, 38 species representing 19 genera are described, including 10 new species, one new genus and one new subgenus: Eirlysella buckhornensis gen. et sp. nov., Shansiella (Oklahomaella) globilineata subgen. et sp. nov., Phymatopleura girtyi, Phymatopleura conica, Worthenia (Yochelsonospira) kuesi, Dictyotomaria turrisbabel, Paragoniozona yanceyi, Spiroscala shwedagoniformis, Peruvispira oklahomaensis, Baylea tenera. Replacement names are Paragoniozona ornata nom. nov. (for Pleurotomaria aspera Girty), Spiroscala quasipulchra nom. nov. (for Euconospira pulchra Batten). The early ontogenetic shells including protoconchs and early teleoconchs are reported in detail for the first time for most taxa. Most species have a protoconch of one whorl as that of living Vetigastropoda. Planktotrophic protoconchs (multi‐whorled larval shells with sinusigera) are reported for Platyzona and Peruvispira; they are therefore placed in the family Goniasmatidae (Caenogastropoda). Repaired shell scars were found in juvenile Pleurotomariida specimens (c. 1 mm), suggesting exposure to predation from an early stage of ontogeny. Pleurotomariida are strongly dominant in surface samples of the Finis Shale (Texas) but in bulk samples using fine mesh‐sizes, dominance is much less pronounced, indicating a change in clade proportion depending on sampling method. The taxonomic richness and abundance of Pleurotomariida seen in these Carboniferous shales have not been reported from post‐Triassic formations.
A new Permian gastropod assemblage from the Roadian (Middle Permian) Khao Khad Formation, Saraburi Group (Lopburi Province, Central Thailand) which is part of the Indochina Terrane, has yielded one of the most diverse Permian gastropod faunas known from Thailand. A total of 44 gastropod species belonging to 30 genera are described herein, including thirteen new species and one new genus. The new genus is Altotomaria. The new species are Bellerophon erawanensis, Biarmeaspira mazaevi, Apachella thailandensis, Gosseletina microstriata, Worthenia humiligrada, Altotomaria reticulata, Yunnania inflata, Trachydomia suwanneeae, Trachyspira eleganta, Heterosubulites longusapertura, Platyzona gradata, Trypanocochlea lopburiensis and Streptacis? khaokhadensis. Most of the species in the studied assemblage represent vetigastropods (35.6%) and caenogastropods (26.7%) and most of the species belong to Late Palaeozoic cosmopolitan genera. The studied faunas come from shallow water carbonates that are rich in fusulinids, followed by gastropods, ostracods, bivalves and brachiopods. The gastropod assemblage from the Khao Khad Formation shares no species with the gastropod assemblages from other Permian formations in Thailand, the Tak Fa Limestone and the Ratburi Limestone. However, it is similar to the Late Permian gastropod faunas from South China of the Palaeo-Tethys, therefore it suggests that the Indochina Terrane was not located far from South China.
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