Abundant new ichthyosaur material has recently been documented in the Slottsmøya Member of the Agardhfjellet Formation from the Svalbard archipelago of Norway. Here we describe a partial skeleton of a new taxon, Janusaurus lundi, that includes much of the skull and representative portions of the postcranium. The new taxon is diagnosed by a suite of cranial character states including a very gracile stapedial shaft, the presence of a dorsal process on the prearticular and autapomorphic postcranial features such as the presence of an interclavicular trough and a conspicuous anterodorsal process of the ilium. The peculiar morphology of the ilia indicates a previously unrecognized degree of morphological variation in the pelvic girdle of ophthalmosaurids. We also present a large species level phylogenetic analysis of ophthalmosaurids including new and undescribed ichthyosaur material from the Upper Jurassic of Svalbard. Our results recover all Svalbard taxa in a single unresolved polytomy nested within Ophthalmosaurinae, which considerably increases the taxonomic composition of this clade. The paleobiogeographical implications of this result suggest the presence of a single clade of Boreal ophthalmosaurid ichthyosaurs that existed during the latest Jurassic, a pattern also reflected in the high degree of endemicity among some Boreal invertebrates, particularly ammonoids. Recent and ongoing descriptions of marine reptiles from the Slottsmøya Member Lagerstätte provide important new data to test hypotheses of marine amniote faunal turnover at the Jurassic-Cretaceous boundary.
In spite of a fossil record spanning over 150 million years, pelvic girdle evolution in Ichthyopterygia is poorly known. Here, we examine pelvic girdle size relationships using quantitative methods and new ophthalmosaurid material from the Slottsmøya Member Lagerstätte of Svalbard, Norway. One of these new specimens, which preserves the most complete ichthyosaur pelvic girdle from the Cretaceous, is described herein as a new taxon, Keilhauia nui gen. et sp. nov. It represents the most complete Berriasian ichthyosaur known and the youngest yet described from the Slottsmøya Member. It is diagnosed on the basis of two autapomorphies from the pelvic girdle, including an ilium that is anteroposteriorly expanded at its dorsal end and an ischiopubis that is shorter or subequal in length to the femur, as well as a unique character combination. The Slottsmøya Member Lagerstätte ichthyosaurs are significant in that they represent a diverse assemblage of ophthalmosaurids that existed immediately preceding and across the Jurassic–Cretaceous boundary. They also exhibit considerable variation in pelvic girdle morphology, and expand the known range in size variation of pelvic girdle elements in the clade.
Summary. Variation in Lateglacial lithic assemblages has led some researchers to sub‐divide the British Later Upper Palaeolithic into two or more typologically distinct phases (Campbell 1977; Jacobi 1988). Until now, however, the chronological relationships between the different phases have been difficult to demonstrate. In this paper we present new dating evidence for two distinct blade technologies of Lateglacial type found in southwest Britain. The first corresponds to the ‘Creswellian’(as defined by Jacobi 1991; Jacobi and Roberts 1992), and is dated to the earlier part of the Lateglacial Interstadial (c. 13–12,000 years BP); the second technology compares with European Federmessergruppen assemblages (literally, ‘penknife group’ industries, as described in Schwabedissen 1954) of the ‘forested’ phase of the Interstadial (c. 12–11,000 years BP). In addition to presenting the new dates, this paper explores the implications of treating the technologies as separate entities and puts forward preliminary ideas on differences in raw material use and other human subsistence behaviour.
Abstract:The Late Jurassic Slottsmøya Member Lagerstätte on Spitsbergen offers a unique opportunity to study the relationships between vertebrate fossil preservation, invertebrate occurrences and depositional environment. In this study, 21 plesiosaurian and 17 ichthyosaur specimens are described with respect to articulation, landing mode, preservation, and possible predation and scavenging. The stratigraphic distribution of marine reptiles in the Slottsmøya Member is analysed, and a correlation between high total organic content, low oxygen levels, few benthic invertebrates and optimal reptile preservation is observed. A new model for 3D preservation of vertebrates in highly compacted organic shales is explained.
The Late Jurassic Slottsmøya Member Lagerstätte on Spitsbergen preserves a diverse array of marine reptiles, including four named taxa of ophthalmosaurid ichthyosaurs. One of these, Palvennia hoybergeti, is based on the single holotype specimen (SVB 1451) with an incomplete skull. A newly discovered specimen (PMO 222.669) with a disarticulated but largely complete skull and anterior postcranium is described, which considerably expands our knowledge of this taxon. Two additional new ophthalmosaurid specimens with pectoral girdles from the same member are described. The taxonomic utility of the ophthalmosaurid pectoral girdle is contentious, and an assessment of seven pectoral girdles from the Slottsmøya Member provides a basis for addressing this question via a 2D landmark principal component analysis of baracromian coracoids. The analysis reveals a taxonomic signal in the coracoids but also highlights the degree of individual variation. Commonly used phylogenetic characters do not fully encapsulate the degree of variation seen in coracoids and in some cases combine analogous features.
ABSTRACT-Colymbosaurus is a genus of long-necked plesiosaurian represented by two valid species: C. megadeirus from the Upper Kimmeridge Clay Formation (Kimmeridgian-Tithonian) of the United Kingdom and C. svalbardensis from the Slottsmøya Member of the Agardhfjellet Formation (Tithonian-Berriasian) of Svalbard, Norway. Due to the lack of complete and articulated skeletons and a near absence of cranial material, Colymbosaurus has been problematic to characterize morphologically. Here, we describe and conduct a phylogenetic analysis on an informative new specimen referable to C. svalbardensis from the Slottsmøya Member, preserving a large portion of the axial and appendicular skeleton. The new material contributes important new osteological data for the species and together with an extensive examination of congeners in British museums, clarifies the diagnostic characters of the genus. We provide two new diagnostic characters of the epipodials for the genus and reevaluate the utility of an anteroposteriorly oriented bisecting ridge on the distal end of the propodials. We also present two new diagnostic features for C. svalbardensis regarding the neural canal and femoral morphology. A phylogenetic analysis recovers a monophyletic and well-supported Colymbosaurus. The new specimen of C. svalbardensis confirms that this species is not synonymous with other described Slottsmøya Member plesiosauroids, demonstrating considerable diversity of the clade at high latitudes close to the Jurassic-Cretaceous boundary.
The palaeontology of the Lower to Middle Triassic succession in Spitsbergen has been studied for more than a century and a half. Our ability to properly interpret the evolutionary and ecological implications of the faunas requires precise stratigraphic control that has only recently become available. Within such a detailed stratigraphic framework, the Spitsbergen fossil material promises to contribute to our understanding of the faunal recovery after the end-Permian mass extinction.
Jaw elements of Omphalosaurus sp. are described from the Early Triassic (Spathian) of Marmierfjellet, Spitsbergen. The elements are from the Grippia and the Lower Saurian niveaus in the Vendomdalen Member of the Vikinghøgda Formation. In the Grippia niveau a bonebed was excavated in 2014-15 and a large number of ichthyopterygian elements were recovered. Together with the omphalosaurian jaw elements a collection of large vertebral centra were recognized as different from the smaller Grippia centra and more than 200 large vertebral centra are referred to Ichthyopterygia indet. and tentatively assigned to regions of the vertebral column. We refrain from further assignment due to the systematic position and the difficulty of defining criteria for recognizing postcranial elements of Omphalosaurus.
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