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The precursory mire of the Middle Pennsylvanian (Bolsovian) Lower Radnice Coal was buried in situ by volcanic ash, preserving the taxonomic composition, spatial distribution, vertical stratification, and synecology of this peat-forming ecosystem in extraordinary detail. Plant fossil remains represent the preeruption vegetation of the swamp, which resulted from accumulation of peat in a high-ash, planar (rheotrophic) mire situated in a narrow palaeovalley containing an active fluvial system. A tuff bed (the Bělka) at the base of the volcaniclastic Whetstone Horizon was exposed in two contiguous excavations over an area of 50 m 2 in the Radnice Basin of western Bohemia, Czech Republic. Twenty-seven morphotaxa were identified, representing 20 whole-plant species with a wide variety of growth forms. The canopy of the peatforming community was dominated by Cordaites borassifolius trees together with the arborescent lycopsid "Lepidodendron" (= Paralycopodites), whereas Lepidophloios cf. acerosus was subdominant. Evidence suggests that the laterally extensive "crowns" of these arborescent lycopsids would have overlapped during the final phase of their life cycles, but differences in the height of tree species resulted in a complex and vertically variable canopy interrupted by randomly distributed gaps. The understorey was dominated by medullosan pteridosperms and marattialean tree ferns, whereas zygopterid ferns and sphenophylls comprised the bulk of the ground cover. In comparison with the canopy, understorey and ground cover species were less abundant and patchier in distribution, with almost complete absence beneath the deep shade of C. borassifolius trees. Lianas that entwined arborescent trees were an important component of the peat-forming forest. Three lyginopterid pteridosperm species along with a sphenophyll had a lianescent habit based on their close association with upright or prone lycopsid trunks and "canopy" branches. Species richness in the swamp superficially appears low. However, considering the small area of excavation, along with the higher diversity known from the same tuff bed in the adjacent, former opencast Ovčín Mine, it appears that species richness in the forest was comparable to some of the less diverse Westphalian peat-forming swamps in the U.S.A. The Lower Radnice mire vegetation was compositionally homogeneous, but had a heterogeneous distribution with patchiness occurring at a very fine scale. The preserved plant assemblage most resembles mires dominated by medullosan pteridosperms and Paralycopodites described from upper Westphalian coal balls in the U.S.A., which were characterised by high diversity in all storeys and involved plants centred in high-ash peat-forming swamps.

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Vegetation is a major driver of fluvial dynamics in modern rivers, but few facies models incorporate its influence. This article partially fills that gap by documenting the stratigraphy, architecture and palaeobotany of the Lower Pennsylvanian Boss Point Formation of Atlantic Canada, which contains some of the Earth's earliest accumulations of large woody debris. Braided-fluvial systems occupied channel belts of varied scale within valleys several tens of metres deep and more than 12 km wide, and their deposits predominantly consist of sandy and gravelly bedforms with subordinate accretionary macroforms, high flow-strength sand sheets and rippled abandonment facies. Discrete accumulations of clastic detritus and woody debris are up to 6 m thick and constitute at least 18% of the in-channel deposits; they represent lags at the base of large and small channels, fills of minor channels and sandy macroforms that developed in central positions in the upper parts of channel fills. Sandstones with roots and other remnants of in situ vegetation demonstrate that vegetated islands were present, and the abundance of discrete channel fills suggests that the formation represents an anabranching, island-braided sandbed river, the earliest example documented to date. Although some sphenopsid and lycopsid remains are present, most woody fragments are derived from cordaitalean trees, and the evolution of this group late in the Mississippian is inferred to have exerted a significant influence on fluvial morphodynamic patterns. The formation records a landscape in which active channel belts alternated with well-drained floodplains colonized by dense, mature forests and local patches of pioneering, disturbance-tolerant vegetation. Lakes and poorly drained floodplains dominated by carbonate and organic deposition, respectively, were also present. A large supply of woody debris triggered channel blockage and avulsion, and active channel margins and islands within the channel belts were initially colonized by pioneer vegetation and subsequently stabilized by large trees. A similar alternation of stable and unstable conditions is observed in modern braided rivers actively influenced by vegetation.

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