Program comprehension is an important cognitive process that inherently eludes direct measurement. Thus, researchers are struggling with providing suitable programming languages, tools, or coding conventions to support developers in their everyday work. In this paper, we explore whether functional magnetic resonance imaging (fMRI), which is well established in cognitive neuroscience, is feasible to more directly measure program comprehension. In a controlled experiment, we observed 17 participants inside an fMRI scanner while they were comprehending short source-code snippets, which we contrasted with locating syntax errors. We found a clear, distinct activation pattern of five brain regions, which are related to working memory, attention, and language processing-all processes that fit well to our understanding of program comprehension. Our results encourage us and, hopefully, other researchers to use fMRI in future studies to measure program comprehension and, in the long run, answer questions, such as: Can we predict whether someone will be an excellent programmer? How effective are new languages and tools for program understanding? How should we train developers?

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It is widely accepted that the perception of human voices is supported by neural structures located along the superior temporal sulci. However, there is an ongoing discussion to what extent the activations found in fMRI studies are evoked by the vocal features themselves or are the result of phonetic processing. To show that the temporal lobes are indeed engaged in voice processing, short utterances spoken by famous and unknown people were presented to healthy young participants whose task it was to identify the familiar speakers. In two event-related fMRI experiments, the temporal lobes were found to differentiate between familiar and unfamiliar voices such that named voices elicited higher BOLD signal intensities than unfamiliar voices. Yet, the temporal cortices did not only discriminate between familiar and unfamiliar voices. Experiment 2, which required overtly spoken responses and allowed to distinguish between four familiarity grades, revealed that there was a fine-grained differentiation between all of these familiarity levels with higher familiarity being associated with larger BOLD signal amplitudes. Finally, we observed a gradual response change such that the BOLD signal differences between unfamiliar and highly familiar voices increased with the distance of an area from the transverse temporal gyri, especially towards the anterior temporal cortex and the middle temporal gyri. Therefore, the results suggest that (the anterior and non-superior portions of) the temporal lobes participate in voice-specific processing independent from phonetic components also involved in spoken speech material.