Two new genera and species of Tritylodontidae, Shartegodon altai and Nuurtherium baruunensis, are described and compared with other known tritylodontids. The new taxa are represented by two partial skulls and, in the case of Nuurtherium, postcranial elements. They were collected from the lower part of the Ulan Malgait Sequence (Upper Jurassic) in the Shar Teg locality, Gobi-Altai Aimag, southwestern Mongolia. The upper postcanine teeth of these two genera have unique cusp formulae that differentiate them from other tritylodontids. As has been suggested for most tritylodontids in general, these two new genera may have been herbivorous.

The San Diego Formation, Pico Formation, Careaga Sandstone, and Foxen Mudstone of southern California are thought to be late Pliocene to early Pleistocene; however, numerical ages have not been determined. Following assessment of diagenetic alteration via multiple methods including scanning electron microscopy (SEM), X-ray diffraction (XRD), and minor elemental concentrations, we attempted to use strontium isotope stratigraphy to assign numerical ages. Using aragonitic fossils, we obtained ages of 2.0−1.85 Ma for the Careaga Sandstone and 2.0−1.75 Ma for the uppermost Foxen Mudstone, consistent with biostratigraphic work suggesting a Gelasian age for the Careaga Sandstone. Isotope ratios for aragonitic and calcitic fossils from the Pico Formation were poorly constrained, with the exception of one bed yielding ages of 5.1−4.3 Ma. Isotope ratios from the San Diego Formation were also inconsistent within beds, with the exception of two isolated outcrops that yielded ages of 5.0−4.5 Ma and 4.5−2.8 Ma, respectively. The age estimates for the Pico and San Diego Formations are older than most ages inferred from biostratigraphy. Noting that some aragonitic specimens from the San Diego Formation yielded isotope ratios indicative of ages as old as 19.4 Ma, we propose that some outcrops have been affected by diagenesis caused by groundwater flow through proximal granitic rocks and input from detrital sediment. Although we recommend that strontium isotope results for the Pico and San Diego Formations be interpreted with caution, the ages of the uppermost Foxen Mudstone and Careaga Sandstone can be confidently placed within the early Pleistocene.

The Cretaceous-Paleogene (KPg) boundary, one of Earth's five major extinction events, occurred just before the appearance of Placentalia in the fossil record. The Gobi Desert, Mongolia and the Western Interior of North America have important fossil mammals occurring just before and after the KPg boundary (e.g. Prodiacodon, Deltatheridium) that have yet to be phylogenetically tested in a character-rich context with molecular data. We present here phylogenetic analyses of >6000 newly scored anatomical observations drawn from six untested fossils and added to the largest existing morphological matrix for mammals. These data are combined with sequence data from 27 nuclear genes. Results show the existence of a new eutherian sister clade to Placentalia, which we name and characterize. The extinct clade Leptictidae is part of this placental sister clade, indicating that the sister clade survived the KPg event to co-exist in ancient ecosystems during the Paleogene radiation of placentals. Analysing the Cretaceous metatherian Deltatheridium in this character-rich context reveals it is a member of Marsupialia, a finding that extends the minimum age of Marsupialia before the KPg boundary. Numerous shared-derived features from multiple anatomical systems support the assignment of Deltatheridium to Marsupialia. Computed tomography scans of exquisite new specimens better document the marsupial-like dental replacement pattern of Deltatheridium. The new placental sister clade has both Asian and North American species, and is ancestrally characterized by shared derived features such as a hind limb modified for saltatorial locomotion.