Comparison of plate convergence with the timing and magnitude of upper crustal shortening in collisional orogens indicates both shortening deficits (200–1700 km) and significant (10–40%) plate deceleration during collision, the cause(s) for which remains debated. The Greater Caucasus Mountains, which result from postcollisional Cenozoic closure of a relict Mesozoic back‐arc basin on the northern margin of the Arabia‐Eurasia collision zone, help reconcile these debates. Here we use U‐Pb detrital zircon provenance data and the regional geology of the Caucasus to investigate the width of the now‐consumed Mesozoic back‐arc basin and its closure history. The provenance data record distinct southern and northern provenance domains that persisted until at least the Miocene. Maximum basin width was likely ~350–400 km. We propose that closure of the back‐arc basin initiated at ~35 Ma, coincident with initial (soft) Arabia‐Eurasia collision along the Bitlis‐Zagros suture, eventually leading to ~5 Ma (hard) collision between the Lesser Caucasus arc and the Scythian platform to form the Greater Caucasus Mountains. Final basin closure triggered deceleration of plate convergence and tectonic reorganization throughout the collision. Postcollisional subduction of such small (102–103 km wide) relict ocean basins can account for both shortening deficits and delays in plate deceleration by accommodating convergence via subduction/underthrusting, although such shortening is easily missed if it occurs along structures hidden within flysch/slate belts. Relict basin closure is likely typical in continental collisions in which the colliding margins are either irregularly shaped or rimmed by extensive back‐arc basins and fringing arcs, such as those in the modern South Pacific.

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The Greater Caucasus orogen on the southern margin of Eurasia is hypothesized to be a young collisional system and may present an opportunity to probe the structural, sedimentary and geodynamic effects of continental collision. We present detrital zircon U-Pb age data from the Caucasus region that constrain changes in sediment routing and source exposure during the late Cenozoic convergence and collision between the Greater Caucasus orogen and the Lesser Caucasus, an arc terrane on the lower plate of the system. During Oligocene to Middle Miocene time, following the initiation of deformation within the Greater Caucasus, marine sandstones and shales were deposited between the Greater and Lesser Caucasus, and detrital zircon age data suggest no mixing of Greater Caucasus and Lesser Caucasus detritus. During Middle to Late Miocene time, Greater Caucasus detritus was deposited onto the Lesser Caucasus basin margin, and terrestrial, largely conglomeratic, sedimentation began between the Greater and Lesser Caucasus. Around 5.3 Ma, upper plate exhumation rates increased and shortening migrated to pro-and retro-wedge fold-thrust belts, coinciding with the initiation of foreland basin erosion. Sediment composition, provenance and structural data from the orogen together suggest the existence of a wide (230-280 km) marine basin that was progressively closed during Oligocene to Late Miocene time, probably by subduction/lithospheric underthrusting beneath the Greater Caucasus, followed by initiation of collision between the Lesser Caucasus arc terrane and the Greater Caucasus in Late Miocene to Pliocene time. The pace of the transition from hypothesized subduction to collision in the Caucasus is consistent with predictions from numerical modeling for a system with moderate convergence rates (<13 mm/yr) and hot lower plate continental lithosphere. Basement crystallization histories implied by our detrital zircon age data suggest the presence of two pre-Jurassic sutures between stable Eurasia and the Lesser Caucasus, which likely guided later deformation.

Deformation within orogenic plateaus functions to establish a dynamic equilibrium between tectonic boundary stresses and plateau gravitational potential energy. Temporal changes in deformation kinematics record perturbations to boundary stresses or internal plateau processes, such as lithospheric foundering. We integrate new mapping, field observations, and geochronologic ages with published data to document complex late Cenozoic upper crustal deformation in the region of the Antofalla depression, a ∼125 km long, sublinear basin within the southern Puna orogenic plateau, Argentina. The juxtaposition of stratal ages across the depression requires >900 m vertical offset on a surface‐breaking fault. Regional geologic structure, basin geomorphology, and our observation of a breccia paralleling the depression margin suggest formation of the depression by normal faulting. We interpret published stratigraphic logs to suggest that the depression formed between ca. 16 and 11 Ma following Andean shortening. Folded Late Miocene to Quaternary strata on the eastern depression margin indicate that extension ended and shortening resumed before present, revealing toggling between extensional and contractional kinematic regimes. The kinematic evolution of the Antofalla depression contrasts with the rest of the southern Puna plateau, which underwent shortening until latest Miocene to Quaternary time, followed by extension and strike‐slip deformation. Taken together, the spatial and temporal variations in late Cenozoic deformation of the southern Puna plateau are inconsistent with mechanisms that would affect the entire orogen, such as slowing convergence, but are compatible with lithospheric foundering.