“…Our understanding of colony organisation and stability in the eusocial insects has been significantly improved by the application of network analyses (e.g., Charbonneau et al, 2013;Fewell, 2003;Jeanson, 2012;Naug, 2009). However, other insect species, many of which show levels of sociality approaching those of the eusocial insects (see Costa, 2006), have been almost entirely overlooked (but see Formica et al, 2012;and Fisher, Rodríguez-Muñoz, & Tregenza, 2016b, for rare examples). In our study, we therefore chose social network analysis as a tool to improve our understanding of insect social associations.…”