“…These rely on evidence of occupancy and activity such as scratch marks, scats, citizen sightings, and bellows [3,20,22,24,25]. In contrast, the application of direct techniques, such as diurnal strip-width [10,26,27] and line-transects [10], nocturnal line-transects [3], and emerging remotely piloted aircraft system thermal imaging (RPAS, also known as drones) technologies [8,[28][29][30], require surveyors to physically sight koalas and this is preferable for density and population estimates [3].…”