“…Seedling recruitment is sensitive to the effect of competition and therefore depends on the presence of gaps in vegetation cover (Amiaud and Touzard, 2004;During and van Tooren, 1990;Ho¨lzel and Otte, 2004;Kitajima and Tilman, 1996;Sˇpacˇkova´et al, 1998;Zamfir, 2000;Zobel et al, 2000). Gaps emerge as a result of small-scale disturbances, for instance from plant deaths, substrate movement or animal activity such as trampling or burrowing (Edwards and Crawley, 1999;Jentsch and Beyschlag, 2003;Pickett and White, 1985;van Tooren, 1988). Diaspores that colonize gaps are derived from several sources, seeds present in the persistent seed bank, seeds produced in current year by adjacent vegetation, or seeds actively relocated by animals to the gaps over longer distances (Bullock et al, 1994;Edwards and Crawley, 1999;Kalamees and Zobel, 2002;Pakeman et al, 1998).…”