Today, most global drylands are located between ∼20° and 40°N, whereas Asian inland drylands, one of the most climate sensitive and vulnerable regions in the world (Lioubimtseva & Henebry, 2009), span from northwestern China to middle Asia at higher latitudes. In particular, central Asia (approximately 40-50°N, 70-105°E) hosts steppe-desert biomes and is characterized by summer-dominated low precipitation (Barbolini et al., 2020). Drylands in central Asia have a very long and complex evolutionary history during the Cenozoic (Caves & Chamberlain, 2018;Lu et al., 2019), accompanied by ancient biomes characterized by a predominance of shrubs, forest, and ferns rather than herbs (