“…In contrast, association with TP, an indicator of nutrient condition, was opposite among haplotypes. This may be due to the intraspecific variation in traits relating to growth and phosphorus-use efficiency, and physiological responses to nutrient limitation (Hartnett, 2019;Sherman et al, 2017Sherman et al, , 2020, as was found experimentally among JPN1 genotypes living in Japan (Tian et al, 2019).We also analysed the effect of the dipteran Chaoborus, whose larva is a predator of Daphnia and whose abundance indicates the abundance of planktivorous fish, because Chaoborus is also preyed on by fish (Sweetman & Smol, 2006). The directions of the effects of Chaoborus also differed among haplotypes, indicating the possibility that predation by this invertebrate predator (Chaoborus larvae) and planktivorous fish on Daphnia influenced the haplotypes in different ways.…”