Upper Cretaceous Foraminifera Murgeina Apula (Luperto Sinni, 1968): A Methusalem and Cenomanian-Turonian Boundary Survivor Taxon

Abstract

The benthic foraminifera Murgeina apula (Luperto Sinni), originally described as Nummofallotia apula from the Upper Cretaceous of southern Italy, represents a common taxon in the middle-upper Cenomanian Sarvak Formation of southwestern Iran as well as the Campanian-early Maastrichtian of Croatia. The specific identity of the Iranian and Italian as well as Croatian specimens is confirmed by equivalent biometric data and identical test structure. The agglutinated wall (inner part) of M. apula excludes its assignment to the porcelaneous Nummofallotia Barrier & Neumann. Nummofallotia is currently placed in the family Meandropsinidae Henson, while for Murgeina with its bilamellar wall inclusion in the Ventrolaminidae Weynschenk is favoured. Nummofallotia cenomana Luperto Sinnni, described from the Cenomanian of southern Italy and lacking any hard facts for species discrimination (e.g., dimensions, inner structure) is herein considered tentatively a junior synonym of Murgeina apula requiring re-examination of the type-material. M. apula represents another survivor of the Cenomanian–Turonian boundary extinction event and, with a long stratigraphic range from the middle Cenomanian to the (early) Maastrichtian, is similar to another Methusalem taxon such as Moncharmontia apenninica (De Castro) with a comparable range.