“…We obtained 55 observations for heterosexual tandems (6, 13, 14, 8, and 14 for colonies A, B, C, E, and F, respectively), which included observations of separation search by 29 leader-females and 26 follower-males. Similarly, we observed 46 female-female same-sex tandems (2,10,13,8,13), with 22 leader-female and 24 followerfemale observations of separation search, and 56 male-male same-sex tandems (6,12,14,8,16), with 29 leader-male and 27 follower-male observations of separation search. We extracted the coordinates of termite movements from all videos using the video-tracking system UMATracker (54).…”