“…Many problems of dietary and feeding habits for the diversely shaped buccal organs remain essentially unsolved, even for most extant cephalopods (Nesis, 1986(Nesis, , 1987Lehmann, 1988). Of particular interest are the aptychus-type mandibles (including anaptychi) present in many but by no means all Mesozoic ammonoids, and the rhynchaptychus-type mandibles of the Lytoceratina (Lehmann et al, 1980). A consensus appears to be that, among the former, the uncalcified anaptychi functioned mainly as feeding organs, i.e., as lower mandibles or, perhaps, an elastic "pump" during suction feeding (Seilacher, 1993); the thickly calcified aptychi (Lamellaptychus, Laevaptychus, Punctaptychus) functioned primarily as opercula (Schindewolf, 1958; Lehmann and Kulicki, 1990;Seilacher, 1993).…”