“…Terminal sclereids have no clear phylogenetic delimitations (Malaviya, 1976; Rao & Bhupal, 1973; Rao & Chin, 1966; Tomlinson, 1959) and originate from parenchyma cells that elongate irregularly into branches invading intercellular spaces and pushing between the walls of neighbouring cells (Evert, 2006; Griffith, 1968). As they are frequently observed in close association with terminal tracheids and veins, terminal sclereids have long been thought to have an extra‐vascular role in water transport (Foster, 1956; Rao & Bhupal, 1973; Rao & Mody, 1961); despite this, their contributions to leaf hydraulics are still being resolved (Heide‐Jorgensen, 1990; Losada et al, 2020). Such extra‐vascular leaf structures have been suggested as adaptations to increase water dispersal from xylem to client cells within the leaf lamina, thereby improving hydraulic efficiency (Brodribb & Holbrook, 2005; Brodribb, 2015; Buckley, John, Scoffoni, & Sack, 2015).…”