Typhlocharis Dieck, 1869 (Coleoptera: Carabidae, Anillini): a new species from the Iberian Peninsula, with notes about its relationships and the evolution of the diecki species group

Abstract

A new species of the genus Typhlocharis Dieck (Coleoptera: Carabidae, Anillini), T. martini, described from the upper Segura River (province of Albacete, southeast Spain), is included in the diecki group characterised by a series of 4 +3 setae in the lateral umbilical series and a reduced number of preapical and apical dentiform projections of the elytron. A key to the eight species of the diecki species group is included. The relationships of the new taxon and the hypotheses about the evolutionary history of Typhlocharis and the diecki species group are discussed. ResumenSe describe una nueva especie del género Typhlocharis Dieck (Coleoptera, Trechidae, Anillini), T. martini, procedente de la cuenca alta del río Segura (Albacete, sureste de España). La nueva especie forma parte del grupo diecki que se caracteriza por tener una serie umbílica de 4+3 sedas en el élitro y un número reducido de dientes en las zonas subapical y apical del élitro. Se incluye una clave de las especies del grupo diecki, y se discuten las relaciones del nuevo taxon y las hipótesis acerca de la historia evolutiva de los Typhlocharis y del grupo diecki.

Two new species of <i>Typhlocharis</i> Dieck, 1869 (Coleoptera, Carabidae) from Portugal: description and notes on the related species

“…The hind trochanters of males are ovoid elongate, not thorn-shaped as happens in T. carmenae and T. farinosae. The form of the spermathecae of T. lunai, T. besucheti Vigna-Taglianti, 1972, T. carmenae, T. farinosae, T. gonzaloi Ortuño, 2005 and T. martini Andújar, Lencina & Serrano, 2008 are si-milar (spheroid), while in the new species it is barrel shaped. The number of apical gonocoxite setae is one like in T. lunai and T. farinosae (2 in T. besucheti, T. carmenae, T. gonzaloi and T. martini).…”

“…So, the present distribution of Typhlocharis may be dated from 25-30 MY onwards. Some comments on the origin and evolution of this genus can be found also in the recent work of Andújar et al (2008).…”

“…Taking into account the identification key given by Andújar et al (2008) for the diecki group species, the new species can enter in this key, with some modifications, as follows:…”

“…Within the subtribe Anillina and according to the catalogue of the ground beetles of the Iberian Peninsula (Serrano, 2003) plus some more recent works (Ortuño, 2005;Serrano et al, 2005;Serrano & Aguiar, 2006a, 2006bAndújar et al, 2008) Typhlocharis are distributed throughout the Iberian Peninsula (Europe), Morocco and Tunisia (North Africa) (Jeannel, 1963). Representatives of this genus have been found in the vestiges of the Lusitanian, the Lionigurian, the Betic Riffian and the Numidian Massifs and the genus is considered a very old one (Jeanne, 1973).…”

“…According to Zaballos & Ruiz-Tapiador (1997) the adults belonging to the species of the diecki and outereloi groups are recognized by the following combination of characters: diecki group, a) umbilicate series of elytra with four setae in the anterior group and three setae in the posterior group (4+3) and b) apical edge of elytron with no more than two teeth; outereloi group, a) umbilicate series of elytra with four setae in the anterior group and one-two setae in the posterior group (4+1 or 4+2) and b) apical edge of elytron with no more than two teeth. The eight known species of the diecki Group were found in Spain, Tunisia and Portugal (Andújar et al, 2008). On the other hand, the fifteen known species of the outereloi group can be found in several regions of Portugal and Spain (e.g.…”