“…Gesnerioideae is itself subdivided into five tribes, of which Beslerieae and Gesnerieae account for most of the species. Gesnerieae includes the species rich genus Columnea L., in which new species continue being described, particularly from Colombia and Ecuador (Amaya-Márquez et al, 2015; Clark et al, 2021; Smith et al, 2013b; Tobar et al, 2022).…”