“…Despite these predictions, there is substantial variability in the speed of treeline response to warming in mountains around the world (Camarero et al, 2017). This variation may be due to many factors, such as varying geomorphology, past human disturbance, lags in population dynamics, dispersal limitation, and biological interactions (competition, herbivory; Grace, Berninger, & Nagy, 2002;Harsch, Hulme, McGlone, & Duncan, 2009;Holtmeier, 2003;Holtmeier & Broll, 2017;Speed, Austrheim, Hester, & Mysterud, 2010). Variability in treeline advance hints that in addition to warming temperature, there are other factors influencing treeline advance, most of which remain unexplored.…”