Nectar secreted by isolated nectaries floated on sucrose solution was of a higher total molar concentration than the medium. However, under steady-state conditions, the 8ucrose concentration was lower than that of the medium, the nectar containing glucose and fructose in addition to sucrose_ When mannitol was added to the medium the osmotic pressure of the nectar exceeded that of the medium although the total molar sugar concentration of the nectar could be somewhat less_The osmotic pressure of phloem exudate (13 atm) was found to be less than that of nectar produced in 8itu (19 atm).Various models to account for nectar production are examined.