The Mars Science Laboratory heat shield was designed to withstand a fully turbulent heat pulse using information from ground testing and computational analysis on a preflight design trajectory. Instrumentation on the flight heat shield measured in-depth temperatures to permit reconstruction of the surface heating. The data indicate that boundary-layer transition occurred at five of seven measurement locations before peak heating. Data oscillations at three pressure measurement locations may also indicate transition. This paper presents the heat shield temperature and pressure data, possible explanations for the timing of boundary-layer transition, and a comparison of reconstructed and computational heating on the actual trajectory. A smooth-wall boundary-layer Reynolds number that was used to predict transition is compared with observed transition at various heat shield locations. A single transition Reynolds number criterion does not uniformly explain the timing of boundary-layer transition observed during flight. A roughness-based Reynolds number suggests that transition due to discrete or distributed roughness elements occurred. However, the distributed roughness height from acreage heat shield material would have needed to be larger than expected. The instrumentation confirmed the predicted location of maximum turbulent heat flux near the lee-side shoulder. The reconstructed heat flux at that location is bounded by smooth-wall turbulent convective heating, indicating that a significant augmentation due to surface roughness did not occur. Turbulent heating on the downstream side of the heat shield nose exceeded smooth-wall convective levels, assuming a supercatalytic surface, indicating that roughness may have augmented heating. The stagnation region heating also exceeded calculated convective heating; the cause of elevated heating may be attributed to a combination of shocklayer radiation and a heating augmentation of unknown origin that was also evident in ground test data.m 2 ∕s D = aeroshell diameter, m H r = recovery enthalpy, J∕kg h = altitude, km h w = wall enthalpy, J∕kg k = roughness height, m k s = equivalent sand-grain roughness height, m k = roughness augmentation Reynolds number, ρ w U τ k s ∕μ w M = Mach number m = entry system mass, kg p = pressure, Earth atm; one Earth atm is equal to 101,325 Pa Q = integrated heat load, ∫ q dt, J∕cm 2 q = heat flux, W∕cm 2 q ∞ = freestream dynamic pressure, 1∕2ρ ∞ V 2 ∞ , Pa Re k;k = roughness Reynolds number, ρ k u k k∕μ k Re θ = momentum thickness Reynolds number, ρ e u e θ∕μ e Re ∞ = freestream Reynolds number, ρ ∞ V ∞ D∕μ ∞ St = Stanton number, q w ∕ρ ∞ V ∞ H ∞ − H w T = temperature,°C t = time from atmospheric interface, s U τ = shear velocity, τ w ∕ρ w p , m∕s V = velocity relative to atmosphere, km∕s α = angle of attack, deg Subscripts D = aeroshell diameter e = boundary-layer edge, where H e is equal to 0.99H ∞ i = atmospheric interface lam = laminar trim = aerodynamic trim condition turb = turbulent w = wall w, k = rough wall w, 0 = smooth wall ∞ = fr...