“…bat guano (Saiz-Jimenez and Hermosin, 1999;Barton and Jurado, 2007;Cuezva et al, 2009;Pasić et al, 2010;Vanderwolf et al, 2013). Caves may also represent heterogeneous systems because of the nature and surface properties of the various internal rock layers, the architecture and connectivity of the cave (Cañveras et al, 2001;Nehme, 2013), as well as airflow and climatic exchanges with the outside atmosphere (Russell, 1998;Banerjee and Joshi, 2013). Outside conditions impact most on the entrance zone of caves (Banerjee and Joshi, 2013), and thus climatic fluctuations within caves depend on the distance to the entrance, as well as epikarst thickness (Cuezva et al, 2009).…”