“…Although the original definition of the T-OAE was based on the presence of a lithostratigraphic marker (Jenkyns, 1985), the development of chemostratigraphy demonstrated that the T-OAE is associated with a negative C isotopic anomaly documented in marine carbonates and organic matter as well as in terrestrial organic matter including fossil wood and specific organic compounds (Jenkyns and Clayton, 1986;Hesselbo et al, 2000Hesselbo et al, , 2007Schouten et al, 2000;Jenkyns et al, 2002;Emmanuel et al, 2006;Van Breugel et al, 2006;Al-Suwaidi et al, 2010;Caruthers et al, 2011;Izumi et al, 2012;Kafousia et al, 2014;Reolid, 2014;Xu et al, 2017;Them et al, 2017;Fantasia et al, 2018). As shown by Fantasia et al (2018), such a negative C isotopic anomaly might have resulted from volcanogenic CO 2 , thermogenic methane associated with metamorphism, and dissociation of marine or terrestrial clathrates.…”