“…These levels are linked with the spiral direction of the sepals (Endress, 1999) so that the result is, strictly speaking, floral asymmetry. Such cases were reported from Ericaceae (personal observation), Polemoniaceae (Grant & Grant, 1965;Mabberley, 2000), Gentianaceae (Thiv & Kadereit, 2002a), Rubiaceae (Robbrecht, 1988), Boraginaceae (Taroda & Gibbs, 1986), Convolvulaceae (Wagner, 1989;Deroin, 1996), and Solanaceae (Robyns, 1930;Hunziker, 2001). These families are all in asterids I (lamiids), except for Ericaceae and Polemoniaceae, which are in the more isolated Ericales (basal asterids).…”