“…Habitat-specific effects on signal evolution have been studied predominantly in visual signals in fish (Boughman, 2001;Endler, 1991Endler, , 1992Seehausen et al, 1997) and birds (Marchetti, 1993), and in the acoustic (air-borne) signals in insects (Larsen and Michelsen, 1983;Romer, 1998), birds (Richards and Wiley, 1980;Ryan and Brenowitz, 1985;Wiley, 1991), and frogs (Ryan et al, 1990;Ryan and Wilczynski, 1991). Signals in most modalities, such as visual and auditory, are adapted to a single medium (e.g.…”