“…This is due in part to the large innate variability that exists in the morphology of mycelia grown in submerged cultures (Metz et al, 1981), which makes its characterization difficult. Quantitative studies include the work of Roels et al (1974), Metz et al (1979), Kim and Yoo (1992), Liu and Yu (1993), Fatile (1985), Olsvik et al (1993), Olsvik and Kristiansen (1994), Mohseni and Allen (1995), Tucker and Thomas (1993), and Tucker (1994). Only the last five of these used image-analysis methods for rapid and accurate characterization of mycelial morphology-methods that greatly extend the amount and type of data that can be obtained.…”