“…Of course, we are by no means suggesting that all observed variations in tropical vegetation structure should be explainable on the simple basis of the CWAK hypothesis. For example, there are numerous cases where poor drainage may be the reason for savanna vegetation types under conditions otherwise suitable for forest (Cole, 1960(Cole, , 1986Haase and Beck, 1989;Cronje et al, 2008;Mantlana et al, 2008;Cochrane and Cochrane, 2010), and it also seems reasonable to speculate that in some cases woody savanna tree extent could be constrained by supra-low calcium concentrations in the subsoil (Cochrane, 1989). Also, with sodium capable of replacing potassium in many of its physiological functions when available at moderate concentrations (Wakeel et al, 2011;Kronzucker et al, 2013), of considerable interest in terms of refinement of the CWAK hypothesis is the existence of very high biomass stands of Colophospermum mopane (J.Kirk ex Benth.)…”