Phylloceratina and Lytoceratina annn onoids in the Middle Jurassic of the Iberian Chain (E. Spain) represent less than I % of the whole of Ann nonoidea. There are two intervals, however, in which their occurrence is remarkably constant: within the Upper Bajocian and at the LowerlMiddle Callovian boundary. These two dispersal episodes of Phylloceratina and Lytoceratina into the Iberian Basin are regarded to reflect changes in their palaeoecological and taphonomical behaviour, as a consequence of regional sea-level changes during the Middle Jurassic. A relative rise during the Late Bajocian favoured the immigration of juvenile phyllocerataceans. Phylloceras and Adabofoloceras immigrations gave rise to monospecific assemblages, where they soon died. They did not breed or ontogenically develop in this basin. In contrast, phyllocerataceans recorded at the LowerlMiddle Callovian boundary constitute poly specific assemblages, dominated by adult individuals. These Callovian assemblages were formed by nekroplanktonic drift, related to a relative sea-level fall and general homogenization of the shelf bottom, hence favouring the concentration of remains of organisms from more open marine and oceanic areas.