“…500, i.e., in the rough-turbulent flow regime. However, in natural channels (and gravel-bed streams in particular) one cannot assume a uniform grain size distribution and equal mobility as in Shields' experiments, as the relative grain size (i.e., the ratio between the grain size of interest, hereafter called D i , and the median grain size D 50 of the channel bed) affects incipient motion thresholds because of hiding and protrusion effects (Komar, 1987a;Gob et al, 2010). Consequently, differences between critical motion thresholds of large and small particles in nonuniform grain size distributions are reduced, a process that needs to be considered when studying boulders on a bed of smaller gravels.…”