“…It has been widely demonstrated that the absorption, transmission, and transformation of light energy by photosynthetic apparatus are carried out on pigmented protein complexes with a certain molecular arrangement and spatial conformation (Ai, Bie, Yuan, Liu, & Bo, ), and the process includes four high‐abundance multisubunit membrane integrin complexes, namely PSI (Photosystem I), PSII (Photosystem II), Cyt b6f (cytochrome b6f complex) and ATPase (ATP synthase), which contain 70 proteins in total (Dekker & Boekema, ; Peng, ; Y. L. Sun et al, ; Wijk, ). In general, ROS are produced in PSI under stress, but it affects both PSI and PSII, leading to damage to the structure and function of thylakoid membrane proteins (Hideg, Spetea, & Vass, ; Rochaix, ; Tjus, Scheller, Andersson, & Møller, ).…”