“…Observations and measurements were made on herbarium material at LY, NOU (Bruy et al 2021; Bruy 2022), and P (Le Bras et al 2017), as well as on specimen images available on line at https://www.recolnat.org. These images, as well as photos of living plants, were measured using the Annotate-on tool (version 1.9.56) made available by the RECOLNAT infrastructure (downloadable from https://www.recolnat.org/fr/annotate).…”