The desire to propagate plants with useful and predictable characteristics vegetatively has been a major concern since the beginning of agriculture. Clonal propagation of apple varieties by grafting onto rootstocks dates back to ancient times (72). A range of techniques for vegetative propagation have been developed over the years, including advances in layering, rooting cuttings, grafting and budding, and the relatively recent approach of micropropagation (31). Vegetative propagation is costly and inefficient compared to seed propagation. For crops with short generation time, repetitive inbreeding or hybrid crosses of inbred lines allows the propagation of nearly genetically uniform populations by sexually produced seed (1). However, generation time in perennial tree crops, including Malus spp., is too long to make this approach practical. As a result, sexually produced seedling populations are highly variable. For this reason, the limited numbers of apomictic species among perennials constitute extremely valuable exceptions, allowing efficient propagation of genetically uniform populations from seed.