“…To determine whether vp8 affects internal tissue as well as the leaf epidermis, double mutants were constructed between vp8and Ragged leavesl (Rgl). Rgl is a dominant mutation that causes cytolysis of mesophyll cells with an indirect effect on epidermal cells (Mericle, 1950). Rgl expression first becomes visible at approximately the same time as other adult traits, is delayed by the heterochronic Teopod mutations, and is unaffected by the epidermisspecific heterochronic mutation, g115 (Bongard-Pierce et aL, 1996;Evans et aL, 1994).…”