“…This has revealed that sex ratio of the resident population at this time can favour males, e.g., J. lagostoma (Hartnoll et al, 2006b;Hartnoll et al, 2009;Hartnoll et al, 2010). The sex ratio of crabs during the migration is variable; it can be skewed towards males at the beginning, e.g., G. ruricola (Hartnoll et al, 2007), G. natalis (Hicks, 1985), and D. hirtipes (Foale, 1999), but be close to unity during the migration itself, e.g., J. lagostoma (Hartnoll et al, 2009;Hartnoll et al, 2010), and later be dominated by females, e.g., G. natalis (Hicks, 1985), especially at the shoreline, e.g., J. lagostoma (Hartnoll et al, 2006b;Hartnoll et al, 2009;Hartnoll et al, 2010). This change in sex ratio during the migration period has been explained in J. lagostoma by the fact that a greater proportion of females compared to males migrate (or participate in the migration for longer periods) and that as expected a greater proportion of females reach the shoreline to deposit their fertilised eggs into the ocean (Hartnoll et al, 2009).…”