“…Current theory borrows from dyadic animal contest theory, for example, Hawk‐Dove evolutionary game theory (Maynard Smith, 1982) and theory of warfare, for example, Lanchester's law of attrition (Lanchester, 1914). These theories often assume that groups act as single entities during conflicts, or that groups are formed of identical individuals (Adams & Mesterton‐Gibbons, 2003; Fearon, 1995; Johnson & Toft, 2014; Rusch & Gavrilets, 2016; Sherratt & Mesterton‐Gibbons, 2013), although some recent theoretical work recognizes heterogeneity of groups (Bornstein, 2003; Gavrilets, 2015; Gavrilets & Fortunato, 2014; Pandit, Pradhan, Balashov, & Van Schaik, 2016). Empirical work highlights that individuals from the same group respond differently to intergroup conflicts—several studies have shown how different classes of individuals contribute to conflicts, including differences between males and females, differences across dominance rank, and differences between those with and without offspring in the group (Arseneau, Taucher, van Schaik, & Willems, 2015; Boydston, Morelli, & Holekamp, 2001; Kitchen & Beehner, 2007; Mares, Young, & Clutton‐Brock, 2012; Meunier, Molina‐Vila, & Perry, 2012; Muller & Mitani, 2002; Thompson et al, 2020; Van Belle, Garber, Estrada, & Di Fiore, 2014; Van Belle & Scarry, 2015).…”