“…The area of karst landform in China is 3.4 million km 2 , covering more than one-third of the total land area, and there are more than half a million karst caves in China (Chen, 2006;Chen, Zhu, & Dai, 2005;Ran & Chen, 1998;Zhang & Zhu, 2012). Surveys on cave fungal diversity have been rare in China, with checklists produced for a few caves (Hsu & Agoramoorthy, 2001;Ma et al, 2015;Man et al, 2015;Wang et al, 2010Wang et al, , 2011Zhang et al, 2017). Few systematic investigations has been reported using molecular tools for the determination of cave fungal communities (Carmichael et al, 2015;Cloutier et al, 2017;Man, 2017;Pfendler et al, 2018).…”