“…The existence of gradients, or clines, of gene frequencies between spatially separate populations can be substantiated by differences in measured amounts of selection, selection coefficients, in the different environments. However, when migration is enlisted to explain clines, the assumptions which need to be made concerning differential selection may be grossly simplified (Hiorns & Harrison, 1978). When such simpler assumptions are made, migration can even be adduced as necessary to the maintenance of these clines over time.…”