This paper reports findings from an 18-month qualitative study that followed the experiences of nine teacher residents, their site professors, site coordinators, clinical teachers, and principals in three Professional Learning Schools (PLSes). The study examined the tensions that emerged as teacher preparation theory intersected with the context-bound realities of daily life in schools and the political constraints that diminish possibilities for inclusive education. The paper addresses implications for teacher preparation programs by reporting how teacher residents negotiated their understanding of and commitment for inclusive education through three themes: (a) critical reflection required, (b) learning is happening, and (c) troubling behavior. Intepreting these themes has implications for programmatic designs in teacher preparation.