“…Whereas random bivalent formation has been suggested by Govindaswami & Pachytene observations by light microscopy revealed occasional interchromosomal and intrachromosomal pairing in monoploids of rice (Chu, 1967), tomato (Ecochard et al, 1969), maize (Ting, 1966;Ford, 1970;Weber & Alexander, 1972) and barley (Sadasivaiah & Kasha, 1971, 1973. These findings were corroborated by studies based on threedimensional reconstruction of pachytene nuclei by electron microscopy that revealed the existence of synaptonemal complexes (SCs) in monoploids of tomato (Menzel & Price, 1966), maize (Ting, 1973), petunias and snapdragons (Sen, 1970) and barley (Gillies, 1974). However, with this methodology only a limited number of nuclei can be analysed, which prevents a quantitative study of SC formation.…”