“…In later work, he considered that T. acutum resulted from the crossing of two other species because of its morphological characteristics, its habitat and the poor development of its pollen (Duval‐Jouve, 1875). Later studies on this genus confirmed the existence of many hybrids, for instance Rouy (1913), Prat (1931), Simonet (1934, 1935a, b), Östergren (1940a, b), Cauderon (1958, 1966) and Cauderon & Saigne (1961). Cauderon (1958, 1966) reported the frequency of hybridization as well as the presence of many intermediate forms.…”