“…There are other densospore producers, such as Porostrobus (Chaloner, 1962;Leary and Mickle, 1989). However, the consistent co-occurrences of megaspores, such as Zonalesporites from Sporangiostrobus, have aided significantly in separating the taxonomic source of densospores in coals and shales (Grebe, 1966;Corsin et al, 1967;Pierart, 1968;Scott and King, 1981;Bartram, 1987;Wagner, 1989). Given the environmental significance attributed to densospore producers, a major study is needed in which megaspores are considered, and consequent finer taxonomic resolution used in making environmental inferences.…”