©
iForest -Biogeosciences and Forestry
IntroductionCrown characteristics are an important component of growth and yield models. Tree crown research contributes to several key forest ecosystem attributes: biodiversity, productivity, forest management, forest environment, and wildlife (Avery & Burkhart 2002). The crown of a tree has a strong influence on stem shape, as foliage provides carbohydrates for tree growth and development of the whole tree and their vertical distribution influences stem shape (Leites & Robinson 2004, Li & Weiskittel 2010. Crown ratio (CR) is considered as an expression of the tree's photosynthetic potential, and therefore commonly included as a key variable in growth and yield models.Tree stem shape has been commonly modeled using taper models (Muhairwe 1994). Taper models are used to estimate diameter along the bole at any given height, so tree volume can then be determined based on these diameters and corresponding heights. The auxiliary variables used to increase the accuracy of existing taper equations include: (1) crown dimensions; (2) stand and site variables; and (3) upper stem diameter measurements (Trincado & Burkhart 2006). Larson (1963) reported that within the crown, stem diameters at particular heights are generally smaller when compared with trees of the same dimensions but shorter crowns. As a result, tree boles cannot be completely described as a function of bole length and diameter. In attempts to describe tree taper, numerous models of varying complexity have been advanced. In most mathematical models, taper is modeled in terms of dbh (diameter at breast height) and total height. A few researchers have considered using crown variables (e.g., crown length -CL, CR, and crown height -CH) as covariates (Newnham 1992, Leites & Robinson 2004, Jiang et al. 2007) for describing tree profiles because of the relationship between crown and stem form development, but previous studies have shown mixed results on the benefit of adding crown variables in taper models. The main crown variable utilized in taper and volume models was CR (Petersson 1999, Jiang et al. 2007, Li & Weiskittel 2010, Jiang & Liu 2011. However, CL is an interesting variable, which may influence the prediction of diameter and volume in combination with CR (Mäkela 2002). For this reason, some forms of CR and CL functions were incorporated into the tree-stem taper and volume prediction models in this study.For stem taper and volume predictions using regression analysis, an appropriate nonlinear function must first be identified, which is a very difficult task. The main reason that artificial neural network (ANN) applications have received attention is that the methodology is comparable to statistical modeling and ANNs can be seen as a complementary effort (without the restrictive assumption of a particular statistical model) or as an alternative approach to fitting nonlinear models to data. Due to the fact that Neural Networks (NNs) attempt to find the best nonlinear function based on the network's complexity, witho...