“…The patterns recovered here mark 4 broad stages in the evolution of the Central African subclade of the cephus group, all of which are likely tied to the episodic glacial cycles of the Pleistocene which would have alternately opened and closed exchange between Bioko and the African mainland since these landmasses are separated by only 35 km and a sea depth of less than 60 m [Siddall et al, 2003;Ngueutchoua and Giresse, 2010;Wronski et al, 2014]. In stage 1, a proto-C. cephus (common ancestor of C. erythrotis , C. cephus , and C. ascanius ) population became either isolated on Bioko, or semi-isolated in Cameroon north of the Sanaga River during a mid-Pleistocene glacial peak [Grubb, 1990[Grubb, , 2001Tosi, 2017].…”