“…All images © MR-B except bottom row left and middle, © J-CS and top right, © Natalie Forssman paths of small mammals, along with other disturbances they make in the soil, contribute to species richness and diversity, and ecosystem engineering effects (Romero, Gonçalves-Souza, Vieira, & Koricheva, 2014;Root-Bernstein & Ebensperger, 2012). Larger animals that make trails include a number of ungulates, such as blackbuck (Antilope cervicapra) (Baskaran, Ramkumaran, & Karthikeyan, 2016), deer (McCaffery, 1976, bongos (Tragelaphus eurycerus) (Klaus-Hügi, Klaus, Schmid, & König, 1999), guanacos (Lama guanicoe) (MR-B pers. obs.…”