“…At least 60 species are known to produce a variety of sounds during courtship, aggression, and prey detection through stridulation, grinding, rasping, and rapidly contracting specialized sonic muscles that beat the swimbladder like a drum (reviewed in Zelick et al 1999). Among freshwater fishes, otophysans (Johnston and Johnson 2000b), gobiids (Lugli et al 1996), percids (Johnston and Johnson 2000a), cichlids (Amorim et al, 2003), and centrarchids (Ballantyne and Colgan 1978a, b, c) use sound in some manner. To date, no one has asked whether sound is part of the umwelt of stickleback fishes (Gasterosteidae).…”