“…Ecological differences across geologic regions and soil types result in diverse strategies among species (Sponchiado et al, 1989; Rao et al, 1996); species from well‐drained soils have long primary roots (e.g., Voorhees et al, 1975), and those from resource‐rich environments display a proliferation of fine roots (e.g., Pinno and Wilson, 2013). Within species, individual root traits (such as root length and diameter) respond plastically to changes in soil edaphic conditions such as soil compaction (Chen et al, 2014), texture (Hamer et al, 2016; Kulmatiski et al, 2017), moisture (e.g., Bell and Sultan, 1999), and nutrient availability (e.g., Zhang and Forde, 2000; Chevalier et al, 2003; Ferguson et al, 2016), and populations vary in these ecological responses. The effect of ecological soil history on the responses of root or shoot modules across environments remains largely an open question (Wright and Westoby, 2000; Pinno and Wilson, 2013; Larson and Funk, 2016).…”