“…However, at least in the Iberian Peninsula, predators that occa sionally catch amphibians and/or squamates are considered opportunistic, and their accumulations are qualitatively and quantitatively representative of the immediate environment surrounding the cave. At Gran Dolina, it has been shown that the primary agents of small-mammal deposition are owls; secondary agents are mammalian carnivores, self-entries, water flows, mudflows, and other agents (Fernandez-Jalvo and Andrews, 1992;Fernandez-Jalvo, 1996). In the case of amphibians, previous study by Pinto L10na and Andrews (1999) have suggested that remains of 1D3/4 and TD5 come from predation by small carnivores; however, of the material studied here, only a few elements show such strong evidence of digestion similar to that made by carnivores (Blain, 2005;Blain et aI., 2008a).…”