“…Both SCR and SCL3 act to repress middle cortex division, as single mutants of either results in precocious middle cortex formation [25,27]. Recently, a predicted transcriptional co-regulator, SEUSS (SEU), was identified as a positive regulator of SHR , SCR and SCL3 , and prevents middle cortex formation by keeping SHR levels high [28]. Although SEU has been shown to be a negative co-regulator in flower development, in middle cortex formation it can directly bind to upstream regions of SHR , SCR and SCL3 [28–30].…”