The order Cypriniformes is the most diverse group within the superorder Ostariophysi and has over 4,200 species, and is natively distributed on all continents except for Antarctic, South America and Australia. The family Cyprinidae is the largest freshwater fish family in Cypriniformes and includes over 3,000 species grouped in about 370 genera (Nelson, Grande, & Wilson, 2016). Interestingly, the order includes the smallest known vertebrate species with a maximum body size of 9 mm and other species can range up to 3 m (Kottelat, Britz, Hui, & Witte, 2006; Froese & Pauly, 2019). Species inhabit an incredible array of habitats including streams, springs, lakes and caves, and the group includes species that are only found in acidic bogs (pH = 3-4) of SouthEast Asia. With species inhabiting a great variety of habitats comes a broad array of adaptations in their ecologies, morphologies, behaviours, colorations and many other features. Thus, this group represents an outstanding lineage with which to conduct a vast array of evolutionary studies. However, historically the evolutionary/phylogenetic relationships in some groups of the vast majority of species have never been